四稜草属(Schnabelia)系统位置问题的探讨

陈介

Chen Cheih

Received:1900-01-01 Revised:1900-01-01 Online:1964-01-18 Published:1964-01-18
Contact: Chen Cheih

Abstract:

The purpose of the present paper is to discuss the systematic position of the genus

Schnabelia originally proposed by H. Handel-Mazzatti in 1921, who considered it as be-

longing to the family Verbenaceae, a point which he further emphasized in 1936. How-

ever, in his paper on the Chinese verbenaceous plants (1932), Prof. Pei Chien thought

the genus should be removed from Verbenaceae into Labiatae and allied it with such

genera as Ajuga and Teucrium on the ground that it also has deeply lobed ovary. Since

then botanists on Chinese plants, such as Prof. Y. Z. Sun, F. C. How, etc. have always

considered Schnabelia Hand.-Mazz. as a genus of Labiatae.

The present writer has recently studied the genus, comparing its morphology of

ovary, calyx lobes, pedicels, pollen grains, etc. with a number of verbenaceous genera,

and comes to the conclusion that the genus should belong to Verbenaceae as H. Handel-

Mazzatti first proposed and that it is closely allied to the genus Caryopteris, especially

C. nepetaefolia (Benth.) Maxim. Moreover, he considers that the deeply lobed ovary

is not at all a good character even for generic separation, for several genera in Ver-

benaceae, such as Caryopteris, Clerodendron, just like Teucrium and Ajuga of Labiatae,

also have deeply lobed ovaries.

From the ample herbarium material, the present writer discovered for the first time

that the genus Schnabelia has two types of flowers, cleistogamous and chasmogamous.

The original founder of the genus did not know this peculiar floral character, as not

mention of it was made in the generic diagnosis. Chienodoxa Sun (1951) proves to be

the present genus, which is based upon the cleistogamous type of flowers of an allied or

perhaps the same species.

陈介. 四稜草属(Schnabelia)系统位置问题的探讨[J]. 中国科学院大学学报.

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