欢迎访问中国科学院大学学报，今天是
RSS English

中国科学院大学学报

• •    下一篇

肋柱花属的系统研究

刘尚武,何廷农   

  • 收稿日期:1900-01-01 修回日期:1900-01-01 出版日期:1992-08-10 发布日期:1992-08-10

Systematic Study on Lomatogonium A. Br. (Gentianaceae)

Liu Shang-Wu, Ho Ting-Nung   

  • Received:1900-01-01 Revised:1900-01-01 Online:1992-08-10 Published:1992-08-10

摘要：

本文对肋柱花属的属下分类、系统发育和地理分布等方面进行了深入研究。文用分支系统学的方法和原理，用计算机PAUP程序处理，获得了几个最简约的支序图。  肋柱花属属于龙胆亚族辐状花冠群，在这群中，论亲缘关系它与辐花属最近，与獐牙菜属次之，而与黄秦艽属关系较远。  獐牙菜属在进化程度上较肋柱花属低，因此它被选为肋柱花属的外类群。 经过支序分析，肋柱花属的18个种根据Hennig的“共近裔性原则”可组合为三个组，其中肉质根茎组为较原始的组，肋柱花组为中级进化水平的组，合萼组是进化程度最高的组。 肋柱花属是北温带分布型的属，分布于亚洲、欧洲及北美洲，直达北极。从种的地理分布型分析，表明秦岭一横断山区是本属的起源与分化中心。 随文报道了一个新组、一个新种和一个新变种。  研究了全部种类的命名模式。

关键词: 肋柱花属, 獐牙菜属, 辐花属, 黄秦艽属, 龙胆属, 系统发育, 分支系统学

Abstract:

The present paper deals with the infrageneric classification, phylogeny and geographic distribution of the genus Lomatogonium.
    A cladistic analysis was undertaken to establish the taxa and to evaluate the relationships between the taxa. The PAUP computer program was used in this analysis. The most parsimonious tree (Cladogram) of the rotate-corolla group of
subtribe Gentianinae shows that Lomatogonium is closely related to Lomatogoniopsis and Swertia, but distantly to Veratrilla. Among them, Swertia is more primitive  than Lomatogonium and hence Sect. Swertia was selected as the outgroup to polarize the character states of ingroup (Lomatogonium). A data matrix of 29 charaters of Lomatogonium was made for constructing the cladogram. Two most parsimonious trees were formed one of which, with the lowest f value, was at last selected as a shortest tree. In this tree 18 species fall into three groups, i.e. Sect. Sarcorhizoma, Sect. Lomatogonium and Sect. Pleurogynella. The former comes at a lower level with more plesiomorphies while the latter at a higher level with
more apomorphies. Lomatogonium is distributed in the northern temperate zone. However, 16 species are centred in Asia and two extend to Europe, or further to the Arctic region, but none has been found from Africa, Australia and South  erica. The analysis of distribution pattern of species shows that the Qinling-Hengduan Mountain region is both the frequence and diversity centers of Lomatogonium. From the cladogram of Lomatogonium (Fig. 5 ), L. perenne appears to occupy the most plesiomorphic node. This is an indication that it is the extant species closest to the ancestral form and it also implies that the ancestral species may reside in the habitat of this species (the Qinlin-Hengduan Mountain region). On the other hand, a  umber of species of Swertia Sect. Swertia also occur in this region today, which indicates that the Qinlin-Hengduan Mountain region may well be the original center of Lomatogonium. From the distribution pattern of L. rotatum, it can be concluded that the time of the origin dates back at least before the Pliocene. After emergence, this genus had first developed and dispersed in the original center and adjacent region, then diverged into two lineages. One gave rise to the widespread species (northern temperate distribution species L.
carinthiacum and L. rotatum), and the other formed the Himalayan species.
A taxonomic revision of the whole genus Lomatogonium is presented.  In this paper, one new section (Sect. Sarcorhizoma), one new species (L. zhongdianense S. W. Liu et T. N. Ho) and one new variety (L. forrestii var. densiflorum S. W. Liu et  T. N. Ho) are described. The key to the species is given. Type studies are made for all the taxa.

Key words: Lomatogonium, Swertia, Lomatogoniopsis, Veratrilla, Gentiana, Phylogeny, Cladistics

首页 | 期刊介绍 | 编委会 | 投稿指南 | 期刊浏览 | 期刊订阅 | 联系我们

版权所有 © 《中国科学院大学学报》编辑部

本系统由北京玛格泰克科技发展有限公司设计开发

技术支持：support@magtech.com.cn

访问统计

总访问量： 今日访问： 当前在线：

联系我们

通讯地址：北京玉泉路19号（甲）中国科学院大学科研楼410

联系电话:（010）88256013