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中国科学院大学学报 ›› 2002, Vol. 40 ›› Issue (5): 433-443.DOI: 10.7523/j.issn.2095-6134.2002.5.007

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囊状嵩草(莎草科)及相关类群的分类

张树仁   

  • 收稿日期:1900-01-01 修回日期:1900-01-01 发布日期:2002-09-10
  • 通讯作者: 张树仁

Classification of Kobresia fragilis C. B. Clarke (Cyperaceae) and related taxa

ZHANG Shu-Ren   

  • Received:1900-01-01 Revised:1900-01-01 Published:2002-09-10
  • Contact: ZHANG Shu-Ren

摘要:

 根据外部形态学和小坚果表皮的微形态学资料,研究了囊状嵩草及其相关类群的分类。在这一
群植物中,有13个相关的学名。根据以前学者用于区别这些植物的形态性状,无法清楚地将它们划分
开。因此作者深入地研究了有关植物的89号200多份标本,重新评价了形态性状的分类学意义。为了
更深入地理解这类植物的分类,应用扫描电子显微镜对其小坚果表皮的微形态进行了研究。根据研究
的结果,确认了囊状嵩草Kobresia fragilis和弧形嵩草K.curvata,其他名称做为囊状嵩草的异名,K.clar-
keana、K.clarkeana var.megalantha和K.curticeps var.gyirongensis被处理为囊状嵩草的新异名。弧形嵩
草从形态学和微形态学两个方面都明显不同于囊状嵩草。研究中还发现囊状嵩草的一号标本采自甘肃
的碌曲,是其分布的省级新记录。

关键词: 囊状蒿草, 弧形嵩草, 分类, 形态学, 微形态学

Abstract:

The classification of Kobresia fragilis C. B. Clarke and related taxa were studied based
on gross morphology and nutlet epidermal micromorphology. About 13 names have been involved in
the species group. They could not be clearly defined using the morphological characters employed by
previous authors. Thus, more than 200 sheets of 89 specimens of related taxa were studied thorough-
ly, and the taxonomic significance of morpbological characters was re-evaluated. In order to further
understand the classification of K. fragilis and its allies, micromorphology of nutlet epidermis of
these taxa was observed under scanning electron microscope. Based on the results, K. curvata C.
B. Clarke and K. fragilis were recognized. Other names were treated as synonymies of K. fragi-
lis. K. clarkeana (Kükenthal) Kükenthal, K. clarkeana var. megalantha Kükenthal and K.
curticeps (C. B. Clarke) Kükenthal var. gyirongensis Y. C. Yang were reduced to new synony-
mies of K. fragilis. K. curvata was distinct from K. fragilis on both morphological and micromor-
phological basis. One specimen of K. fragilis collected from Luqu, Gansu, China, represents a
new distribution record of this species in Gansu.

Key words: Kobresia fragilis, K. curvata, Classification, Morphology, Micromorphology